Toxodontidae
| Toxodontidae Temporal range: Late Oligocene–Holocene PreЄ Є O S D C P T J K Pg N | |
|---|---|
 | |
Mixotoxodon larensis | |
Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | †Notoungulata |
| Suborder: | †Toxodonta |
| Family: | †Toxodontidae Owen, 1845 |
| Subfamilies and Genera | |
†Haplodontheriinae
†Nesodontinae
†Toxodontinae
| |
Xotodon sp. skull at the Museum für Naturkunde, Berlin
Toxodontidae is an extinct family of notoungulate mammals known from the Oligocene to the Holocene (5,000 BP) of South America,[1] with one genus, Mixotoxodon, also known from the Pleistocene of Central America and southwestern North America (Texas).[2] They somewhat resembled rhinoceroses, and had teeth with high crowns and open roots, suggesting that they often fed on tough pampas grass.[3] However, isotopic analyses have led to the conclusion that the most recent forms were grazing and browsing generalists.[4]
Taxonomy
The endemic notoungulate and litoptern ungulates of South America have been shown by studies of collagen and mitochondrial DNA sequences to be a sister group to the perissodactyls.[5][6][7]
In 2014, a study identifying a new species of toxodontid resolved the families phylogenetic relations. The below cladogram was found by the study:[8]
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References
^ Turvey, Samuel T. (2009-05-28). Holocene Extinctions. OUP Oxford. ISBN 9780191579981..mw-parser-output cite.citation{font-style:inherit}.mw-parser-output q{quotes:"""""""'""'"}.mw-parser-output code.cs1-code{color:inherit;background:inherit;border:inherit;padding:inherit}.mw-parser-output .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Lock-green.svg/9px-Lock-green.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-lock-limited a,.mw-parser-output .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Lock-gray-alt-2.svg/9px-Lock-gray-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/thumb/a/aa/Lock-red-alt-2.svg/9px-Lock-red-alt-2.svg.png")no-repeat;background-position:right .1em center}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration{color:#555}.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration span{border-bottom:1px dotted;cursor:help}.mw-parser-output .cs1-hidden-error{display:none;font-size:100%}.mw-parser-output .cs1-visible-error{font-size:100%}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration,.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-right{padding-right:0.2em}
^ E. Lundelius, et al. 2013. The first occurrence of a toxodont (Mammalia, Notoungulata) in the United States. Journal of Vertebrate Paleontology, Vol 33, No 1, pp. 229–232 DOI:10.1080/02724634.2012.711405
^ Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 253. ISBN 1-84028-152-9.
^ MacFadden, Bruce J. (September 2005). "Diet and habitat of toxodont megaherbivores (Mammalia, Notoungulata) from the late Quaternary of South and Central America". Quaternary Research. Elsevier. 64 (2): 113–124. Bibcode:2005QuRes..64..113M. doi:10.1016/j.yqres.2005.05.003.
^ Welker, F.; Collins, M. J.; Thomas, J. A.; Wadsley, M.; Brace, S.; Cappellini, E.; Turvey, S. T.; Reguero, M.; Gelfo, J. N.; Kramarz, A.; Burger, J.; Thomas-Oates, J.; Ashford, D. A.; Ashton, P. D.; Rowsell, K.; Porter, D. M.; Kessler, B.; Fischer, R.; Baessmann, C.; Kaspar, S.; Olsen, J. V.; Kiley, P.; Elliott, J. A.; Kelstrup, C. D.; Mullin, V.; Hofreiter, M.; Willerslev, E.; Hublin, J.-J.; Orlando, L.; Barnes, I.; MacPhee, R. D. E. (2015-03-18). "Ancient proteins resolve the evolutionary history of Darwin's South American ungulates". Nature. 522: 81–84. Bibcode:2015Natur.522...81W. doi:10.1038/nature14249. ISSN 0028-0836. PMID 25799987.
^ Buckley, M. (2015-04-01). "Ancient collagen reveals evolutionary history of the endemic South American 'ungulates'". Proceedings of the Royal Society B: Biological Sciences. 282 (1806): 20142671–20142671. doi:10.1098/rspb.2014.2671. PMC 4426609.
^ Westbury, M.; Baleka, S.; Barlow, A.; Hartmann, S.; Paijmans, J. L. A.; Kramarz, A.; Forasiepi, A. M.; Bond, M.; Gelfo, J. N.; Reguero, M. A.; López-Mendoza, P.; Taglioretti, M.; Scaglia, F.; Rinderknecht, A.; Jones, W.; Mena, F.; Billet, G.; de Muizon, C.; Aguilar, J. L.; MacPhee, R. D. E.; Hofreiter, M. (2017-06-27). "A mitogenomic timetree for Darwin's enigmatic South American mammal Macrauchenia patachonica". Nature Communications. 8: 15951. Bibcode:2017NatCo...815951W. doi:10.1038/ncomms15951.
^ Forasiepi, A. A. M.; Cerdeño, E.; Bond, M.; Schmidt, G. I.; Naipauer, M.; Straehl, F. R.; Martinelli, A. N. G.; Garrido, A. C.; Schmitz, M. D.; Crowley, J. L. (2014). "New toxodontid (Notoungulata) from the Early Miocene of Mendoza, Argentina". Paläontologische Zeitschrift. doi:10.1007/s12542-014-0233-5.
- McKenna, Malcolm C., and Bell, Susan K. 1997. Classification of Mammals Above the Species Level. Columbia University Press, New York, 631 pp.
ISBN 0-231-11013-8
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